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Pheugopedius vs. Sphenura (1 Viewer)

Norbert R.

Active member
While updating my files on the Troglodytidae I stumbled over the use of Pheugopedius Cabanis, 1851 (type species Pheugopedius genibarbis Cabanis) for a genus of wrens formerly included within Thryothorus Vieillot, 1816 in H&M 4 as well as HBW/BirdLife Illustr. Checklist.

There is the much older name Sphenura M. H. C. Lichtenstein, 1822 (type species Sphenura coraya = Turdus coraya J. F. Gmelin), which has been used for a subgenus of Thryothorus by Wolters (1980), with Pheugopedius in synonymy.

I assume the use of Pheugopedius follows Mann et al. (2006), who split Thryothorus into several genera, one of which a new one: Cantorchilus, but probably overlooked Sphenura.

Is there any reason to treat Sphenura M. H. C. Lichtenstein, 1822 as unavailable? Any suggestions are welcome.
 
Wolters, H. E. (1980): Die Vogelarten der Erde. Eine systematische Liste mit Verbreitungsangaben sowie deutschen und englischen Namen. Lieferung 6, pp. 401-452 (November 1980). Troglodytidae on pp. 430-433.

Also, Hellmayr (1934; Field Mus. Nat. Hist. Publ., Zool. Ser. 13, pt. 7, pp. 110-295) treated Sphenura M. H. C. Lichtenstein, 1822 (Verz. Vögel Berl. Mus., pp. 7,8) as a valid genus-group name, referring to Mathews (1923) Birds Australia 10, p. 149.

So Sphenura seems to be no nomen oblitum and should probably replace Pheugopedius (?).
 
If Sphenura was used as a valid genus after 1900 in several publications , it can't be considered as a Nomen Oblitum, in theory. However , it may have a reversal of precedence
 
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Have you seen the 1822 work ? (I have not.) My understanding is that Lichtenstein 1822 would have included 4 wholly undescribed species (i.e., species which he himself subsequently described in 1823), plus "Sphenura coraya" without any indication of authority, so the name might presumably also be seen a completely nude at this point.

In 1823, he presented the name as new again, indicated four (!) distinct taxonomic species as "Typus" (not a valid type designation), and included 10 more taxonomic species. In 1835, Rüppell designated Sphenura acaciae Licht. (one of the 10 "added" species) as the type. In 1840, Gray designated Turdus brachypterus Latham (the first of the four species making up the originally indicated "Typus") as the type. Should Rüppell's designation be accepted, Sphenura Licht. 1823 would be a senior synonym of Argya; but, in practice, Gray's designation was long accepted, which made it a senior synonym of Dasyornis.

It's the type of name that you'd prefer not to come across...
 
Also, Hellmayr (1934; Field Mus. Nat. Hist. Publ., Zool. Ser. 13, pt. 7, pp. 110-295) treated Sphenura M. H. C. Lichtenstein, 1822 (Verz. Vögel Berl. Mus., pp. 7,8) as a valid genus-group name, referring to Mathews (1923) Birds Australia 10, p. 149.
He did not, actually.
https://biodiversitylibrary.org/page/2767874
Hellmayr certainly treated this name as an available genus-group name, authored by Lichtenstein 1822 and with Turdus coraya Gmelin as its type (as per Mathews 1923). But this is not what the Code calls treating a name as "valid". A "valid name" is a name that you effectively use as the name of a taxon in your classification.
The generic name that Hellmayr used there was Thryothorus Vieill. 1816; he defined no subgenera within it; he listed Sphenura Licht. 1822 merely as an invalid junior synonym.
This type of treatment would not prevent a reversal of precedence. (But of course, if Wolters used it for a subgenus, this should be enough.)

(Another issue, for a reversal of precedence (= making an unused senior name -- here Sphenura -- the nomen oblitum of some widely and long-used junior nomen protectum -- here Pheugopedius), is that the junior name's usage in the recent published literature must be sufficient to allow it. For names that were recently revived, this may not be as straightforward as it may appear at first sight.)
 
Thank you, Laurent, for your well-founded and interessant thoughts about this tricky case.

So we have Hellmayr (1934) who accepted Mathews's (1923) logic that Sphenura Licht., 1822 is an available name. As he did not use subgenera, he circumvented a decision which of the two names should be used for the species group recently revived as the genus Pheugopedius.
Wolters (1980) was for many decades the first author who accepted Sphenura Licht., 1822 as the valid name for a genus-group. Following Hellmayr, Sphenura, Pheugopedius as well as Thryophilus S. F. Baird, 1864 have been subsumed by all 20th century authorities into Thryothorus. Pheugopedius was already synonymized with Thryothorus by Sharpe (1882, Cat. Birds Brit. Mus. 6; and 1903, Hand-list Genera Species Birds 4), whereas Sharpe only used Sphenura Licht., 1823, which is a homonym of Lichtenstein's (1822) earlier name as well as a synonym of Dasyornis.

Mann et al. (2006, Mol. Phylogen. Evol. 40: 750-759) were the first authors for many decades who questioned the genus Thryothorus as circumscribed by Hellmayr (1934). They did not, however, discuss their decision to apply Pheugopedius instead of Sphenura, although both names were not in use, or almost so, for nearly a century. They certainly overlooked that Wolters (1980) chose Sphenura for a subgenus, probably accepting Mathews' s and Hellmayr's decision that Lichtenstein's name of 1822 is available and nearly 30 years older as the next oldest available name. Perhaps it would be better to follow Wolters (1980), but the decision should be made by SACC (?).
 
Laurent wrote:
Have you seen the 1822 work ? (I have not.)
I think it is here:
http://www.archive.org/stream/abhandlungenderp20akad#page/284/mode/2up/search/Lichtenstein .
Sorry that is the Fish.
The amphibians are on page 237.

Birds?
https://zs.thulb.uni-jena.de/rsc/viewer/jportal_derivate_00218073/Isis_1824_Bd01_761.tif . Which cites Isis?
Coues discusses this in Bulletin of the United States Geological and Geographical Survey Volume 5.
And Bibliographia Zoologiæ Et Geologiæ: A General Catalogue Agassiz.
 
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It should be titled (cf. [here]):

Verzeichniss von Vögeln, Conchylien und Insecten, Doubletten des zoologischen Museums hiesiger Königl. Universität : welche am 29sten Jul. u. folg. Tage d. J. Nachmittags um 3 Uhr im Universitäts-Gebäude durch den Königl. Auctions-Commissarius Bratring gegen gleich baare Bezahlung in kling. Preuss. Courant meistbietend versteigert werden sollen.
 
Thank you Laurent. Verzeichniss von Vögeln, Conchylien und Insecten is the sixth item in a volume containing thirteen pamphlets, bound together subsequent to publication. Binder's title: Zoologisches Museum, Berlin. Catalogues, 1816-1856. A few of the thirteen are in BHL:
1854. https://www.biodiversitylibrary.org/item/247839#page/11/mode/1up . (See Sphenura Lich. on page 30)
1842. https://www.biodiversitylibrary.org/item/92528#page/3/mode/1up .
1823. https://www.biodiversitylibrary.org/item/89675#page/11/mode/1up . (see pg 40 Sphenura new genus with 4 type species https://www.biodiversitylibrary.org/item/89675#page/60/mode/1up .)
The 1822 one is similar to the 1842 one? an advertisement of a sale in an afternoon edition for July 29th 1822 newspaper? (Or the sales date at the auction house was afternoon of 29 July 1822?)

Completely unrelated is a Lichtenstein name Turdus figulus.
It is in the 1816 sales catalogue but NN. In the 1819 sales catalogue NN also both according to Richmond but fully described in 1823. But in the 1818 here on page sixty it has enough not to be nomen nudem?
Turdus figulus Lichtenstein,MHC 1819
Turdus figulus Lichtenstein 1816
Turdus figulus Wied 1821
Turdus figulus Lichtenstein,MHC 1823
https://books.google.com/books?id=8...nd+Insecten,+Doubletten&source=gbs_navlinks_s see page 60.
According to Whittell The literature of Australian birds 1818 is a second edition of 1816.
Yes same language in 1816 and 1818 versions of Turdus figulus.
Page 69 of https://books.google.com/books?id=-...+Universitaet+zu+Berlin&source=gbs_navlinks_s .
 
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From Mathews: “To the Isis 1821, as Beylage No. 1, appears a " Zweites Preisverzeichniss der Doubletten des Zoologischen Museums der Konigl. Universitat zu Berlin" by Lichtenstein. In this List there is quite a large number of names, and it seems that many of the names commonly accepted from the very well-known 1823 List will date back to 1821. We have not collated in detail these Lists, but such a work should be undertaken by those interested in African and American
Birds”
Here is the link to that 1821 Isis: https://books.google.com/books?id=_...letten+des+zoologischen&source=gbs_navlinks_s . It is in
Beylage 3 no 1 search for Zweites Preisverzeichniss it is a few pages after 371.
I looked for Sphenura but did not find it.
 
According to C. Richmond in the 1820 Lichtenstein list :Verz. Vögeln Conch. Ins. Doubl. Zool. Mus
Sphenura had species of
coraya Sphenura, M. H. C. Lichtenstein, Verz. Vögeln Conch. Ins. Doubl. Zool. Mus. Berlin, 1820, 8 [n. n.].
frontalis Sphenura, M. H. C. Lichtenstein, Verz. Vögeln Conch. Ins. Doubl. Zool. Mus. Berlin, 1820, 8 [n. n.].
poliocephala (Tem.)Sphenura, M. H. C. Lichtenstein, Verz. Vögeln Conch. Ins. Doubl. Zool. Mus. Berlin, 1820, 7 [n. n.].
sulphurascens Sphenura, M. H. C. Lichtenstein, Verz. Vögeln Conch. Ins. Doubl. Zool. Mus. Berlin, 1820, 8 [n. n.].
superciliaris (Tem.) Sphenura, M. H. C. Lichtenstein, Verz. Vögeln Conch. Ins. Doubl. Zool. Mus. Berlin, 1820, 8 [n. n.].
Different from 1823 list of four types and S. acaciae. Richmond considered all 1820 names N. N.
http://www.sil.si.edu/DigitalCollections/indexanimalium/TaxonomicNames/IA_Namesearch.cfm .
1823 types of Sphenura Turdus brachypterus Latham, Turdus coraya Gm., Certhia cinnamomia, Gm. Fringilla macroura.
 
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Sphenura use in 1901:
https://books.google.com/books?id=joYUAAAAYAAJ&dq="Sphenura"&source=gbs_navlinks_s .
Richmond's reason for calling the 1820 & 1822 Spheura species names n. n. is Lichtenstein did not list any authorities for these names. But two are from Temminck (Ident by Sundevall 1836 Orn. Syst.)and one Gmelin.
https://www.zoonomen.net/cit/RI/Genera/S/s00545a.jpg .
Sph. frontalis is probably Certhia frontalis Latham 1801 which Lich. used in 1818.
S. sulphurescens is Turdus sulphurescens Licht. used by him in a 1819 list.
 
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There is a pamphlet by Lichtenstein dated 1822 which includes Brasilian birds maybe Sphenura? Lichtenstein, M.H.C. (1822) Verzeichnis der, aus der Sendung des Herrn Bescke im Monat Oktober 1822 für die Sammlung des zoologischen Museums ausgesuchten Säugetiere und Vögel. Zool. Mus., Signatur: S I, Bescke, C.F.C. + Bescke C.H., Historische Bild- u. Schriftgutsammlungen, Bestand, 94 pp.
https://www.mapress.com/j/zt/article/view/zootaxa.4250.1.1 .
https://www.birdforum.net/showthread.php?t=353678&highlight=Bescke .
 
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Sphenura use in 1901:
https://books.google.com/books?id=joYUAAAAYAAJ&dq="Sphenura"&source=gbs_navlinks_s .
Richmond's reason for calling the 1820 & 1822 Spheura species names n. n. is Lichtenstein did not list any authorities for these names. But two are from Temminck (Ident by Sundevall 1836 Orn. Syst.)and one Gmelin.
https://www.zoonomen.net/cit/RI/Genera/S/s00545a.jpg .
Sph. frontalis is probably Certhia frontalis Latham 1801 which Lich. used in 1818.
S. sulphurescens is Turdus sulphurescens Licht. used by him in a 1819 list.

This agrees with Mauersberger's (Annalen f. Ornith. 12, 1988) statemant that the 1822 "Verzeichniss" contains no descriptions of new taxa, and that Lichtenstein provided no names of authors nor the origin of the birds which are offered for auction. If this is correct, than Sphenura Lichtenstein, 1822 is not available according to the current Code, regardless what Mathews and Hellmayr wrote. It follows that Sphenura Lichtenstein, 1823 would have been the correct name for the genus currently called Dasyornis, but the latter is now well established and no reversal of precedence is possible.
 
Chances are that Lichtenstein 1822 attributed the names to himself ('n.' for nobis), because this is apparently what he did with his new combinations. (He cited the original author of the species name only in synonymy.)

The type designation by Rüppell ([here], third footnote) is now dated 1840 (not 1835, as I noted above and as the title page of the work suggests). Gray's 1840 List of the genera of birds is assumed to have been published before April, so his designation of Turdus brachypterus Latham as the type of Sphenura ([here]) may indeed have precedence over Rüppell's designation of Sphenura acaciae Lichtenstein.

Note that, if "no reversal of precedence is possible" (which is clearly the case -- Sphenura has been used as the valid name of the bristlebirds many times, including by Mathews himself, after 1899 (even if possibly not after 1923 -- last apparent use in BHL: Hall 1922 [here])), Sphenura retains its precedence over Dasyornis, despite the latter being well established. This means that under the standard provisions of the Code, if the two names apply to the same taxon, Sphenura must in principle be used. The only way out of this is via the submission of a case to the Commission requesting the suppression of Sphenura.
 
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But ... as Rüppel's (third) foot-note reads "Es sind dieses die unter dem Namen Malurus acaciae und M. squamiceps abgebildeten Vögel. Lichtenstein in seinem Doubletten-Verzeichniss des Berliner Museums. Seite 40, beschrieb den erste als den Typus einer neuen von ihm unter dem Namen Sphenura aufgestellten Gattung." ... doesn't this indicate that it was referring to page 40 in the far more well-known Verzeichniss der Doubletten des Zoologischen Museums der Königl. Universität zu Berlin (nebst Beschreibung vieler bisher unbekannter Arten von Säugethieren, Vögeln, Amphibien und Fischen), of 1823 (here):
SPHENURA.​
Novum genus, intermedium inter Certhias, Sittas, Sylvias, Turdus.
[...]​
.

The (sale catalogue) "Verzeichniss von Vögeln, Conchylien und Insecten, ..." (of 1822) apparently only have 31 pages.

If of any help?

Björn
 
But ... as Rüppel's (third) foot-note reads "Es sind dieses die unter dem Namen Malurus acaciae und M. squamiceps abgebildeten Vögel. Lichtenstein in seinem Doubletten-Verzeichniss des Berliner Museums. Seite 40, beschrieb den erste als den Typus einer neuen von ihm unter dem Namen Sphenura aufgestellten Gattung." ... doesn't this indicate that it was referring to page 40 in the far more well-known Verzeichniss der Doubletten des Zoologischen Museums der Königl. Universität zu Berlin (nebst Beschreibung vieler bisher unbekannter Arten von Säugethieren, Vögeln, Amphibien und Fischen), of 1823 (here):

The (sale catalogue) "Verzeichniss von Vögeln, Conchylien und Insecten, ..." (of 1822) apparently only have 31 pages.
Yes, Rüppell's designation applies to the name taken from the 1823 publication. So does Gray's. In both, the designated type was a species that was not included in 1822.

These designations are potentially important only if (as suggested by Norbert in his last post) the name is deemed nude in the 1822 list, because "Sphenura coraya n." without any description (what I think would/will be found in the 1822 work) cannot "be unambiguously assigned to a nominal species-group taxon or taxa" (ICZN 12.2.5 not fulfilled; contra Mathews). If so, the name must be taken from the 1823 work and, given that the "type designation" there cannot possibly be interpreted as valid, its application will depend on the first subsequent designation as its type of any of the nominal species included in the genus in this work, which was either by Gray 1840, or by Rüppell 1840 -- more likely the former.

(If the name is accepted as available from the 1822 list, its type is fixed originally by monotypy (the so-called 'virtual' monotypy -- only one species denoted by an available name in the OD, even though more were included). If so, no subsequent designation can affect it. The 1823 instance is then either (as suggested by Norbert above) a junior homonym, to which one of these two designations might still apply but which cannot possibly become valid because it is preoccupied by the 1822 name; or (my preference) a mere subsequent use without any standing, incorrectly flagged as new (i.e., Lichtenstein himself did not regard his earlier use of the same name for a subset of the species included in 1823 as having established it, while we now do), which had its type fixed from the start in the 1822 work. Under the latter reading: the two subsequent designations of species not included in 1822 are both simply invalid; there is no such thing as a "Sphenura Licht. 1823", but only a Sphenura Licht. 1822 -- in the original sense (Licht. 1822 and 1823 -- the 5 spp names cited in 1822 were all still included in 1823; I see absolutely no suggestion that Lichtenstein's own concept had changed), sensu Gray 1840 (a misidentification), or sensu Rüppell 1840 (another misidentification).)
 
Now I have seen the relevant pages of Lichtenstein's "Preisverzeichniss" of 1822 (through the courtesy of P. Eckhoff, Zool. Mus. Berlin). All names in that list are nomina nuda, as Mauersberger (1988) already stated. They don't have the "n." (for new combinations) or "N." (for new taxa) that Mathews (1925) claims, nor do they have references on earlier authors (e.g. Linn.Gmel.) or on the origin of the specimens offered for sale. I doubt that Mathews actually ever saw Lichtenstein's 1822 "Preisverzeichniss". Hellmayr and Wolters may have followed him becuse the rarity of that publication prevented them from checking the original source. The 1822 publication by Lichtenstein has no relevance for nomenclature, as all names are nomina nuda; it is even impossible to assume that Sphenura coraya is the same as Gmelin's Turdus coraya. Thus, Pheugopedius is the correct name for the wren genus. More problematic, the Australian bristlebirds should again change their genus-group name from Dasyornis to Sphenura Licht., 1823, as Mathews's claim that Sphenura Licht., 1822 is an older homonym of the 1823 name is wrong!
 
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